Optimized for fast shooting speeds and working in difficult conditions, the AF-S NIKKOR 500mm f/4E FL ED VR is an impressive telephoto designed for Nikon FX-format DSLRs. The sophisticated optical design incorporates two fluorite and three extra-low dispersion elements to control chromatic aberrations and distortions, as well as lessen the overall weight of the lens. Also contributing to improved image quality, both Nano Crystal and Super Integrated Coatings have been applied to lens elements to lessen flare and ghosting for improved contrast, clarity, and color fidelity. Protecting the advanced optical design is a front fluorine-coated meniscus glass element that, when paired with the weather-resistant magnesium-alloy barrel, supports working in inclement conditions.

A Silent Wave Motor AF system produces fast, near-silent focusing performance that suits both stills shooting and movie recording, which can instantly be manually overridden by simply turning the manual focus ring. Further contributing to photographing in fast-paced situations and difficult lighting, VR (Vibration Reduction) image stabilization compensates for the effects of camera shake by up to four shutter speed steps. Additionally, an electromagnetic aperture mechanism is integrated into the lens design to provide greater exposure control stability that is especially beneficial when working with faster continuous shooting rates.